ULtrastructure of the Oocyte Vitelline Envelope Deposition in the Amazonian Fish Pseudotylosurus microps (Teleostei: Belonidae)
Abstract
The development of the vitelline envelope during the oogénesis of the “needlefish" Pseudotylosurus microps was examined by transmission electron microscopy. In the previtellogenic oocytes, the vitelline envelope has a bílaminar structure. The inner electron-dense layer seems to be deposited by the oocyte, the outer, less electron-dense, and granule-fibrillar layer seéms to have an exogenous origin. In the vitellogenin oocytes, the vitelline envelope acquires a trilaminar appearance, The outer layer before mentioned become gradually compacted and electron-dense. In a subsequent step, it becomes central by the addition of an outer third stratus with loose aspect and low electron density. Al the same time, the inner electron-dense layer thickens ínwardly by farming domeshaped protuberances covered by less electron-dense cúpulas. The thickening of the median electron-dense and the outer layer seems to occur by the contribution of material arísing from large electron-dense bodies Incaled in (he intercellular spaces of the follicular epithelium. In the mature vitelline envelope the inner (thickest layer has become probably produced by the oocytes as a result of the apposition of endogenous material over the cúpulas, although the participation of exogenous sources could not be discarded mainly at the final part of it deposition. The median layer seems to be co-produced by the oocyte and exogen sources and the outer layer is exogenous in origin. The large electron-dense bodies intermingled with the follicular cells give a very peculiar feature to the follicles, but its specific function is unknown.